Notes
Notes
[Boxed or purple texts indicate personal notes :] • Heterotrophs: Not capable of manufacturing their
TOPIC 01: INTRODUCTION TO ZOOLOGY own food and must rely on external food sources.
o Autotrophs: Produce their own energy
The Science of Zoology using substances from their environment,
usually sunlight, water, and carbon dioxide.
▪ Most autotrophs use a process
• Zoology is the study of animal life.
called Photosynthesis, where
o Zoon – animals
sunlight is used to turn carbon
o Logos - study
dioxide and water into food (sugar)
• Zoologists strive to understand:
and oxygen.
o The origin of animal diversity.
• Cells lack cell walls
o How animals perform basic life processes.
o Animal lacks cell walls, plants have it.
o How they are able to inhabit various
ecosystems.
Characteristics of Life
The Uses of Principles
Molecules in living organisms are more complex than those
• Principles of modern zoology are derived from: in non-living things, although non-living things also contain
o Laws of physics and chemistry molecules.
o Scientific method
• Because life shares a common evolutionary origin, 1. Chemical Uniqueness – Living systems demonstrate a
principles learned from the study of one group often unique and complex molecular organization.
pertain to other groups as well.
Living organisms assemble large molecules –
macromolecules – that are more complex than molecules
Zoology As Part of Biology found in nonliving matter.
Example: Humans eat rice → body converts it to glucose → Example: Some bacteria become antibiotic-resistant over
glucose becomes ATP → ATP powers body activities. time.
5. Metabolism - Living organisms maintain themselves by 8. Development - All organisms pass through characteristic
acquiring nutrients from their environments. stages in their life cycle.
• Metabolism includes all of the chemical reactions • Development includes characteristic changes an
occurring within an organism. organism passes through from its beginning
• Digestion - breaking down food (usually as a fertilized egg) through adulthood
• Cellular Respiration - converting nutrients into • Metamorphosis - transformation from one life
energy stage to another.
• Synthesis of molecules and structures - o Tadpole to frog
building important molecules and structures o Caterpillar to butterfly
Metabolism has two types of reactions: Growth – measurable increase in size or mass.
Development – changes in form or function that may not
1. Catabolic reactions (Destructive) – breaking down always be measurable.
molecules to release energy
2. Anabolic reactions (Constructive) – building
Example:
complex molecules using energy
> Growth: baby becoming taller
These processes also involve the formation and breakdown > Development: baby learning to walk or speak
of the four macromolecules.
9. Environmental Interaction – Living organisms interact
These reactions include synthesis of the 4 types of with their environments.
macromolecules as well as breakage of bonds to recover the • Ecology is the study of this interaction between
energy stored there. organisms and between organisms and their
environment.
> Metabolism is not only digestion. o This includes how organisms get food,
> It includes both breaking down and building processes. shelter, and survive in ecosystems.
Flow Example: Food → broken down into glucose → glucose Living organisms respond to their surroundings.
converted to ATP (catabolic).
ATP → used to build proteins and molecules (anabolic) Example:
> Plants grow toward sunlight.
6. Reproduction - Living systems can reproduce > Humans wear jackets when it is cold.
themselves. Life does not arise spontaneously but comes
only from prior life, through reproduction. At each level of 10. Movement - Living systems and their parts show precise
the biological hierarchy living forms reproduce to generate and controlled movements arising from within the system.
others like themselves. • Living systems extract energy from their
• Genes replicate themselves forming new genes. environments permitting the initiationof controlled
• Cells divide to produce new cells. movements.
Movements at the cellular level are required for: Perpetual Change - The world and the organisms living in it
are always changing.
• Reproduction
• Growth • Supported by the fossil record.
• Responses to stimuli • The properties of organisms undergo transformation
• Development in multicellular organisms across generations throughout time.
• Theory upon which the remaining 4 are based.
On a larger scale:
> This theory is the foundation for the other four theories.
• Entire populations or species may disperse from one > Life is always changing and evolving.
geographic location to another over time. > Species today are different from species in the past.
o Populations may move or migrate to new
locations over time. Example: Fossils show that ancient horses were much smaller
than modern horses
Movement of nonliving matter:
• Not precisely controlled by the moving objects. Common Descent - All forms of life descended from a
common ancestor through a branching of lineages.
• Often involves external forces.
• Life’s history has the structure of a branching
> Not all movements are visible. evolutionary tree, known as a phylogeny
> Some movements happen inside the body or cells.
• Serves as the basis for our taxonomic classification of
animals
Example:
• Descent with modification.
> Sperm moving toward an egg cell during reproduction.
> Blood circulating in the body. • Supported by molecular work
1. Chemical Uniqueness – living organisms have Example: Humans, monkeys, and apes share a common
complex macromolecules ancestor.
2. Hierarchical Organization – arranged from small to
bigger systems Multiplication of Species - New species are produced by the
splitting and transforming of older species.
3. Regulation – organisms maintain stable internal
conditions (homeostasis) One species can divide into different species over time.
4. Energy Utilization – using energy for life processes
5. Metabolism – all chemical reactions in the body Example: Different species of Darwin’s finches evolved from
6. Reproduction – producing new organisms a common ancestor bird.
7. Evolution – change over generations
8. Development – life cycle changes Gradualism - Large differences result from the accumulation
9. Environmental Interaction – interacting with of small changes over long periods of time.
surroundings
• Occasionally, changes can happen more quickly
10. Movement – controlled movement in organisms
Sometimes, changes may happen faster, but evolution usually
Evolutionary Perspective of Zoology occurs gradually.
Example: Camouflaged insects survive more easily because 2. All organisms are composed of one or more cells.
predators cannot easily see them. • Every living thing is made of cells.
• Some organisms have only one cell, while others have
Adaptation many cells.
• Natural selection explains why organisms are Examples:
constructed to meet the demands of their Bacteria – only one cell (unicellular).
environments. Humans, plants, animals – many cells (multicellular).
• Adaptation results when the most favorable variants
accumulate over evolutionary time.
3. All cells arise from pre-existing cells.
Unity in Diversity • New cells come from cells that already exist.
• Cells divide to produce new cells.
Example: When you get a cut, your body makes new skin cells
to heal it.
Cell Theory
1. Bacteria – simple prokaryotic organisms (example: E. • Organelles are tiny, specialized structures inside a
coli). cell that perform specific tasks necessary for cell
2. Archaea – prokaryotes that often live in extreme survival. Each organelle has a unique function, such
environments (example: hot springs, salty lakes). as producing energy, making proteins, or removing
3. Eukarya – organisms with eukaryotic cells (includes waste.
plants, animals, fungi, and protists).
Organelles are like small organs inside the cell that perform
Some organisms can make their own food through different jobs.
photosynthesis, mainly plants and some protists.
Examples:
Prokaryotic cells - no nucleus or other membrane-bound • Mitochondria – produces energy (ATP).
organelles. • Ribosomes – make proteins.
• Nucleus – contains DNA and controls cell activities.
• Kingdom Archaebacteria
• Kingdom Eubacteria
Components of Eukaryotic Cell
Eukaryotic cells - do have nucleus and membrane-bound
organelles.
The Plasma Membrane The Nucleus
• Kingdom Protista surrounds the cell. The membrane-bound
• Kingdom Fungi organelle found in
• Kingdom Plantae eukaryotic cells
• Kingdom Animalia
Cytoplasm refers to Organelles such as
Animal Cell
the cellular material the mitochondria,
• An animal cell is a type of eukaryotic cell that makes between the cell golgi complex,
membrane and centrioles, and
up the tissues and organs of animals. Unlike plant
nuclear envelope. endoplasmic
cells, animal cells do not have a cell wall or
reticulum
chloroplasts, but they contain specialized structures
Plasma Membrane are found in the
called organelles. cytoplasm.
• It is also frequently called the cell membrane, is a
fundamental biological structure.
• Plasma membrane structure is described using the
fluid mosaic model.
Functions
• Selective permeability
• Protection and barrier
• Cell Signaling
• Transport of Substances
• Structural support
Plant Cells • Cell-to-cell Interaction
• Have a cell wall which gives them structure and Two layers of phospholipid molecules oriented with
protection. hydrophilic heads toward the outside and hydrophobic tails
• Usually have a more rectangular or box-like shape. inside.
• Contain chloroplasts.
• Fluid-like - flexible
Animal Cells
Chloroplast
Phospholipid Bilayer
• Contains chromosomes, which contain DNA. Example: Liver cells have many smooth ER because they
• DNA holds the instructions for making proteins. detoxify chemicals.
Ribosomes = protein-making machines Lysosomes = waste disposal / cleaning system
Cytoskeleton
Microtubules
• Cylindrical, barrel-shaped structures, composed of • Cell membranes are selectively permeable – water
microtubules, that are essential for forming can pass through, but not most solutes.
centrosomes, cilia, and flagella, playing a crucial role • Gases (oxygen & carbon dioxide), urea, lipid soluble
in cell division and organization. solutes can cross the membrane.
o Not everything can pass the membrane.
Made of microtubules. Only:
Helps organize structures during cell division. ▪ small
▪ nonpolar
Cilia & Flagella
▪ or special molecules
• Cilia & flagella are motile extensions of the cell o Big or charged ones need help (proteins)
surface.
• In many single celled organisms they are a source of Diffusion is the movement of molecules from an area of high
locomotion. concentration to an area of low concentration. This tends to
• In multicellular animals they usually sweep material equalize the concentration.
past the fixed cell. • Down the concentration gradient.
• Solutes are molecules (e.g. salt) that are found in a
Structures used for movement (locomotion). solution.
• G1 (Growth 1):
o Cell grows, makes RNA and functional
proteins.
o Example: Proteins needed for cell function are
produced here.
1. Prophase (Prepare) o Haploid (n): Cells that have only 1 set of
• Chromosomes condense enough to be seen (visible) chromosomes (23 in humans).
with a light microscope. ▪ Formed by meiosis.
o Spindle fibers form between the 2 centrioles. ▪ Example: Sperm and egg cells.
o Spindle fibers attach to kinetochores (protein ▪ Function: Ensures that when
structure on chromosomes). fertilization happens, the zygote has
▪ Tip to remember: Think the correct diploid number.
“Prophase = Prepare the
chromosomes and spindle for
action.” • Chromosomes have replicated during interphase
just as in mitosis.
2. Metaphase (Middle)
• Alignment of the chromosomes along center of cell • Meiosis actually consists of two separate divisions.
(metaphase plate). o Meiosis I – serves to separate the two
o Fibers attached to kinetochores on both versions of the chromosome (homologues).
sides of each chromosome. ▪ Separates homologous
chromosomes (one from each
3. Anaphase (Apart) parent).
• Separation of the sister chromatids. ▪ Example: Mom’s chromosome 1
goes to one cell, Dad’s
• Centromere splits apart sister chromatids move
chromosome 1 goes to another.
toward opposite poles.
o Meiosis II – serves to separate the two
o Disassembly of the tubulin subunits shortens
replicas of each version (sister chromatids)
the microtubules.
▪ Separates sister chromatids of
▪ Tip to remember: “Ana = Apart” →
each chromosome.
chromatids move away from each
▪ Example: Each replicated
other.
chromosome splits so each
daughter cell gets one copy.
4. Telophase (Two)
• Re-formation of the nuclei once the chromosomes are
• Because there is only one replication of DNA but two
at opposite poles.
cell divisions, each of the four daughter cells is
o Chromosomes unwind
haploid – has only one set of chromosomes.
▪ Tip to remember: “Telophase =
Two nuclei” → preparing for Result: 4 haploid daughter cells (1 set of chromosomes
cytokinesis. each).
• Only one DNA replication occurs, but two cell
Cytokinesis
divisions happen.
• Cytokinesis – division of the cytoplasm. • These haploid cells are gametes (eggs or sperm).
• Two complete, diploid cells that are identical to the
original cell Extra Notes: Meiosis introduces genetic variation through
• During cytokinesis in animal cells, the cell pinches in crossing over (exchange of DNA between homologous
two. chromosomes) and independent assortment.
• A cleavage furrow produced by microfilaments
deepens until the cell splits Fertilization
Autosomal Linkage
o
• In incomplete dominance, the phenotype of a • Linked genes are located close together on the same
heterozygote appears to be intermediate to, or distinct chromosome.
from, the homozygous dominant and homozygous o They are usually inherited together.
recessive conditions. o They do not follow the rule of independent
assortment!
Dihybrid Cross
• Some traits can be affected by the environment. • Diagram the Punnett square for the offspring between
o Exposure to sunlight affects skin color in the genotypes GgMM (paternal) and ggmm
humans. (maternal). G is dominant for having a widow’s peak
o Nutrition affects height in humans. and M is dominant for having rolled tongue.
o Soil acidity affects color in hydrangea o Genotypic Ratio: 8:8
flowers. ▪ 50% Widow’s peak, rolled tongue
▪ 50% No widow’s peak, rolled
The Chromosomal Theory of Inheritance tongue
Genes
Nitrogenous Bases
DNA
• Gene expression – the use of information in DNA to • Charles Darwin presented evidence that many
direct the production of particular proteins. modern organisms descended from ancestral species
o Transcription – first stage of gene that were different.
expression. A messenger RNA (mRNA) is
Pre-Darwinian Evolutionary Ideas
synthesized from a gene within DNA.
o Translation – second stage – mRNA is used • Early Greek philosophers recorded an early idea of
to direct production of a protein. evolutionary change
DNA Coding • Aristotle recognized fossils as forms of ancient life.
o He developed the Scala Naturae (Scale of
• DNA codes for the sequence of amino acids in a Nature).
protein. ▪ A ranking system from simple to
• A codon is three base-pairs long and is a segment of complex organisms
mRNA that codes for an amino acid. ▪ Example: plants → animals →
humans
Transcription ▪ He believed species were fixed
(they don’t change)
• Messenger RNA (mRNA) transcribes the DNA and • The ancient Greeks didn’t propose an evolutionary
transports it out of the nucleus. mechanism.
Translation o The Greeks did not explain how evolution
happens
• Translation occurs on ribosomes outside the o No mechanism (no process behind the
nucleus. change)
• mRNA attaches to a ribosome and protein synthesis
begins.
• Transfer RNA (tRNA) collects free amino acids from
the cytoplasm and delivers them to the polysome
• Lyell concluded the age of the earth must be millions
of years.
• He stressed the gradual nature of geological changes.
• Lamarck was the first to suggest an explanation for Evolution in Need of a Mechanism
evolution.
o He proposed Inheritance of acquired • Darwin was not the first to have the thought that
characteristics organisms change through time.
o Did not withstand testing o His grandfather, Erasmus Darwin, and
others suggested that life evolves as
A Mechanism for Evolution environments change.
o But a mechanism for that change was
• Darwin presented a mechanism for evolution – needed
natural selection.
Darwin (1809 – 1882)
o Organisms that are in some way more
successful at reproduction will pass on more • Darwin had a lifelong love of nature.
of their genes. • His father wanted him to study medicine.
o Over time the traits responsible for that o This was not what Darwin wanted and he
success will become widespread in the didn’t finish.
population. • After leaving medical school he attended Cambridge
University with the intention of entering the clergy.
• His mentor and botany professor, John Henslow,
recommended him for a position as ship’s naturalist
aboard the Beagle.
• Fossils provide support for the idea that life changes Anatomical Homologies
through time.
o Fossil intermediates • Vertebrate embryos have a tail and pharyngeal
▪ Whales descended from land pouches.
mammals. • These structures develop into different but
▪ Birds descended from one branch homologous structures in adults.
of dinosaurs. o Gills in fishes
• The oldest fossils are of prokaryotes o Part of ears & throat in humans
• Geological time can be measured in sedimentary rock • Genetic Similarities: Species that are closely related
layers. have more similar DNA sequences. Example:
o The Law of Stratigraphy Humans and chimpanzees share over 98% of their
▪ Dates oldest layers at the bottom DNA.
and youngest at the top. Homologies & the Tree of Life
• Radiometric dating methods are based on the decay
of naturally occurring elements into other elements • Darwin’s evolutionary tree of life can explain
homologies.
Examples: o The genetic code is shared by all species
because it goes back deep into the ancestral
• 40K has a half life of 1.3 billion years – meaning half
past.
of the 40K will have decayed to 40Ar and 40Ca. Half
of what remains will decay in the next 1.3 billion
years.
Evolutionary Trends Forces of Evolutionary Change
• The fossil record shows that species arise and go • Evolution occurs when allele frequencies in a
extinct repeatedly throughout geological history. population change over time. The five main forces
• Trends appear in the fossil record – directional that drive this change are:
changes in features or patterns of diversity • Mutation, gene flow, genetic drift, natural selection,
sexual selection.
Common Descent
Natural Selection
• Darwin proposed that all organisms have descended
from a single ancestral form. Natural selection - Organisms that are more adapted to their
• Life history is shown as a branching tree called a environment are more likely to survive and pass on the genes
phylogeny that aided their success. This process causes species to
change and diverge over time.
Speciation
Genetic Drift
• Speciation is the evolutionary process by which new
biological species arise. • The smaller the sample, the greater the chance of
• It occurs when a population of organisms becomes deviation from expected results.
reproductively isolated from its original group, leading o Evolution occurs due to chance events,
to genetic divergence. especially in small populations.
• Which allele was lost is due to random chance.
Why Does Speciation Matter? • Over time, drift tends to reduce genetic variation
through random loss of alleles.
• It increases biodiversity.
• It explains the diversity of life forms we see today. The Bottleneck Effect
• It helps scientists understand evolutionary
relationships. • Sometimes a catastrophic event can severely reduce
• Speciation refers to the formation of new species. the size of a population.
• Defining a species is difficult… o The random assortment of survivors may
o Descent from common ancestral population. have drastically different allele frequencies.
o Ability to interbreed. o Bottleneck effect
o Maintenance of genotypic & phenotypic • The actions of people sometimes cause bottlenecks
cohesion. in other species.
• Reproductive barriers prevent species from o N. California elephant seal population
interbreeding. reduced to 20 100 individuals in the 1890s.
o Variation drastically reduced
Allopatric Speciation
Mutation
• Allopatric derives from “allo” meaning “separate” and
“patria” meaning “country”. • New genes or alleles only result by mutations.
• Over time, genetic differences accumulate, leading to • Mutations are changes in the nucleotide sequence of
a new species. DNA.
• The geographical separation can arise in two ways: • Beneficial mutations of any kind are very rare.
o Vicariant speciation is initiated when • Mutations that alter gene number or sequence are
climatic or geological changes fragment a almost always harmful.
species’ habitat, forming impenetrable
Point Mutations
barriers.
o Founder events occur when a small number • Point mutation – a change in a single base pair.
of individuals disperse to a distant place • Often harmless
where no other members of their species o Much of the DNA does not code for protein
exist. products.
• Happens when a new species evolves within the o Genetic code is redundant.
same geographic area. ▪ CGU, CGA, CGC, CGG all code for
• Often due to genetic mutations, behavioral arginine.
differences, or habitat preferences. • Occasionally significant
o Sickle cell disease.
Parapatric Speciation
• Sexual recombination is a much more common way TOPIC 05: THE REPRODUCTIVE PROCESS
of producing variation in populations.
Reproduction
o Reshuffling of allele combinations already
present in the population is how variation is • Reproduction is one of the fundamental properties of
maintained in populations. life.
o Sexual reproduction rearranges alleles into • Evolution is inextricably linked to reproduction.
fresh combinations every generation. • Two modes of reproduction are recognized:
Sexual Selection o Asexual
o Sexual
• Sexual selection natural selection for mating success.
o May result in sexual dimorphism Asexual vs. Sexual Reproduction
differences between the sexes.
o Secondary sexual characteristics – not • Asexual Reproduction – the production of offspring
directly involved in reproduction. whose genes all come from one parent without the
fusion of egg and sperm.
Intrasexual Selection o Usually diploid eggs are produced by mitosis
which then develop directly.
• Intrasexual selection – selection within the same
sex – results when individuals of one sex are Asexual reproduction means making a new living thing
competing with each other for members of the other without needing two parents.
sex.
o Features that make the male a better fighter One parent can create an offspring that is basically a
or more intimidating to other males would be copy of itself.
favored. • No mating involved
Intersexual Selection • No mixing of genes
• Offspring are usually identical (clones)
• Intersexual selection – mate choice – individuals of
one sex are choosy in selecting a mate. Example: A plant growing a new plant from its stem, or
o Features that make an individual more bacteria splitting into two.
attractive to the opposite sex would be
favored. • Sexual Reproduction – the production of offspring
• Involves one sex (usually females) choosing mates by the fusion of haploid gametes (eggs & sperm)
from the other sex based on certain traits. from two parents to form a diploid zygote (fertilized
• Showiness that results from mate choice can be risky. egg).
• Benefits of finding a mate outweigh potential costs. Sexual reproduction means making a new living thing
Mass Extinctions using two parents.
• Mass extinctions are episodic events where many A male and a female combine their cells to create a
species go extinct at the same time. baby.
o Permian extinction – 225 MYA – half the • Needs two parents
families of shallow-water marine • Genes are mixed from both
invertebrates and 90% of the marine • Offspring are not identical (they’re unique)
invertebrate species went extinct over a few Example: Humans having a baby, or animals reproducing.
million years.
o Cretaceous extinction – 65 MYA – marks Genetic variability is increased by the random combinations
the end of the dinosaurs as well as many of genes from the parents.
other species
• Many possible explanations for mass extinctions have Asexual Reproduction
been suggested. • Bacteria and many protozoa can reproduce by
Binary Fission – separating into two or more
individuals approximately the same size.
o Bacteria and some tiny organisms can • Many invertebrates with both sexual and asexual
reproduce by splitting into two equal parts. modes enjoy the advantages of both.
• Budding is a form of asexual reproduction where new
individuals form as offshoots of a parent. Modes of Reproduction
• The offspring may separate or remain attached to
form colonies. Parthenogenesis
o A small “bud” forms on the parent, grows,
and becomes a new individual. • Parthenogenesis involves the development of an
o It can break off and live on its own, or It can embryo from an unfertilized egg or one where sperm
stay attached and form a group (colony) & egg nuclei did not fuse.
• Freshwater sponges release specialized groups of o Asexual Cloning - Making an exact copy of
cells called Gemmules that can grow into new an organism using just one parent.
individuals. o A female can have offspring without a male.
• Fragmentation results when an organism’s body is • In some animals (Daphnia) the females can produce
broken into several pieces and each piece grows into two types of eggs.
a new organism. o One must be fertilized.
o Regeneration – the regrowth of lost body o One type will develop directly into haploid
parts. adults – parthenogenesis.
▪ Fragmentation = break and make • In many social insects, like honeybees, males
new ones (drones) are haploid and are produced by
▪ Regeneration = grow back what’s parthenogenesis while females (workers & queens)
missing develop from fertilized eggs
• Sea stars can regenerate lost limbs, but only species • Parthenogenesis occurs in vertebrates in some
in the genus Linckia can form new individuals from fishes, amphibians, lizards, and has recently been
broken arms. discovered in snakes .
o After meiosis, the chromosomes are
Asexual Reproduction - Advantages doubled, creating diploid “zygotes”.
o Often mating behavior is required to
• Animals living far from members of their own species stimulate development of offspring.
can reproduce without having to search for a mate.
• Produce numerous offspring quickly – ideal for Hermaphroditism
colonizing a new area.
• Advantageous in a stable, favorable environment • Hermaphroditism occurs when an organism has
both male and female reproductive systems.
because it reproduces a successful genotype
o One organism has both male and female
precisely.
parts.
Sexual Reproduction
Sequential Hermaphroditism
• Generally involves two parents.
• In sequential hermaphroditism, an organism
• Special germ cells unite to form a zygote.
reverses its sex during its lifetime.
• Sexual reproduction recombines parental characters.
o An organism can change from male to
• In haploid asexual organisms’ mutations are
female (or female to male) during its life.
expressed and selected quickly.
• Protandy - male to female e.g. clownfish
• In sexual reproduction a normal gene on the
• Protogyny - female to male. e.g. parrot fish
homologous chromosome may mask a gene
mutation Fertilization
Why do so many animals reproduce sexually rather than • Fertilization – fusion of egg and sperm into a single
asexually? diploid cell, the zygote.
o The joining of male and female cells to form
• The costs of sexual reproduction are greater than
a new organism.
asexual methods:
o External
o More complicated.
o Internal
o Requires more time.
o Uses more energy. External Fertilization
However: • External Fertilization – fertilization takes place
outside the female’s body.
• Sexual organisms produce more novel genotypes to
o A wet environment is required so gametes
survive in times of environmental change.
don’t dry out and so sperm may swim to the
• In crowded habitats, selection is intense and
eggs.
diversity prevents extinction.
• Environmental cues (day length, temperature) or
• On a geological timescale sexual lineages with less
chemical cues may cause a whole population to
variation are prone to extinction.
release gametes at once.
o Increases likelihood of fertilization.
Internal Fertilization • Viviparous – eggs develop in oviduct or uterus,
nourishment from mother.
• Internal Fertilization allows terrestrial animals to o Mammals, some sharks, scorpions.
reproduce away from water.
o Cooperative behavior leading to copulation is Invertebrate Reproductive Systems
required.
• Many insects have separate sexes, internal
Pheromones fertilization and have complex reproductive systems.
o Female crickets use long ovipositors to
• Pheromones are chemical signals released into the deposit eggs.
environment by one organism that influence the
physiology or behavior of members of the same Gamete Production & Delivery
species.
o Effective in very small amounts. • In vertebrates, reproductive systems are similar with a
o Mate attractants. few important variations.
o Nonmammalian vertebrates usually have
Ensuring Survival of Offspring one combined opening, the cloaca
o Most mammals have a uterus where the
• Species with external fertilization produce huge embryo develops
quantities of gametes that result in lots of zygotes.
o Predation on young is high. Female Reproductive System
o Few will survive to reproduce.
• Species with internal fertilization produce fewer • Ovaries are where female gametes, egg cells, are
zygotes, but protect them more from predation. produced.
o Tough eggshells o A follicle contains one egg cell as well as
o Embryo may develop in reproductive tract of follicle cells that nurture the developing egg.
female ⚬ Most/all of the follicles a woman will
o Parental care of eggs & offspring produce have formed before birth.
• Each month from puberty through menopause one
Advantages of Sexual Reproduction follicle ruptures and releases its egg cell – ovulation.
o The corpus luteum forms from the ruptured
• Sexual reproduction has costs including finding
follicle and secretes estrogen and
mates, greater energy cost, reduced proportion of
progesterone to help maintain the uterine
genes passed on to offspring, and slower population
lining during pregnancy. If the egg is not
growth.
fertilized the lining disintegrates.
• However, sexual reproduction increases variability
• After ovulation, the egg leaves the ovary and enters
in the population – important during times of
the oviduct, which it follows to the uterus.
environmental change.
Reproductive Cycles in Female Mammals
Gamete Production & Delivery
• Humans & some other primates have a menstrual
• Gametes (eggs & sperm) are required for sexual cycle while other mammals have an estrous cycle.
reproduction. o In both, ovulation occurs at a time when the
• Usually, gametes are produced in gonads (ovaries & endometrial lining of the uterus is ready for
testes). an embryo to implant.
• Germ cells are set aside early in development. They o If no egg is fertilized, the lining is shed
will produce only gametes. (menstruation) in humans & other primates
and is reabsorbed in other mammals.
Gametogenesis • Female mammals that have estrous cycles may have
more behavioral changes.
• Gametogenesis – the production of gametes. • Estrous cycles may be more closely tied to season
o Spermatogenesis - the biological process of and climate.
sperm cell production in male organisms,
• Females will usually only mate when in estrous – the
occurring in the testes.
time surrounding ovulation.
o Oogenesis – Oogenesis is the biological
process of egg (ovum) production in female Female Reproductive Cycle
organisms, occurring in the ovaries.
• The female reproductive cycle in humans contains
Reproductive Patterns two parts:
o Uterine (menstrual) cycle
• Oviparous – animals that lay eggs.
o Ovarian cycle
o Most invertebrates, many vertebrates
• Ovoviviparous – animals that retain the eggs within
their bodies. Nourishment comes from the egg.
o Some annelids, insects, some fishes,
reptiles.
The Uterine Cycle Key Events in Development
• If the egg is not fertilized, the corpus luteum • Development describes the changes in an organism
disintegrates, and production of estrogen and from its earliest beginnings through maturity.
progesterone drops sharply. This triggers breakdown o Search for commonalities.
of the endometrium – menstruation. • Specialization of cell types occurs as a hierarchy of
developmental decisions.
Male Reproductive System o Cell types arise from conditions created in
preceding stages.
• Testes are where male gametes, sperm cells, are
o Interactions become increasingly restrictive
produced.
o Testes contain the seminiferous tubules Fertilization
where sperm are formed.
o Sperm production can’t occur at normal body • Fertilization is the initial event in development in
temperature in mammals, so the testes are sexual reproduction.
held outside the body abdominal cavity in the o Union of male and female gametes
scrotum. o Provides for recombination of paternal and
• After leaving the testes, sperm pass through the maternal genes.
epididymis where they become motile and gain the ▪ Restores the diploid number.
ability to fertilize an egg. o Activates the egg to begin development.
• Sperm leave the body through the vas deferens and
urethra. Cleavage
• In males, the principle sex hormones are androgens, • Cleavage – rapid cell divisions following fertilization.
including testosterone. o Very little growth occurs.
o Produced mainly by Leydig cells in the o Each cell called a blastomere.
testes. o Morula – solid ball of cells. First 5-7
o Responsible for secondary sexual divisions.
characteristics.
o Important determinants of behavior in Polarity
vertebrates.
▪ Sex drive • The eggs and zygotes of many animals (not
▪ Aggression mammals) have a definite polarity.
▪ Calling in birds & frogs • The polarity is defined by the distribution of yolk.
• The vegetal pole has the most yolk and the animal
Reproductive Cycles pole has the least.
• Males produce sperm continuously, whereas females Blastula
only release one or a few eggs at certain intervals.
• A fluid filled cavity, the blastocoel, forms within the
TOPIC 06: PRINCIPLES OF DEVELOPMENT embryo – a hollow ball of cells now called a blastula.
Gastrulation
Early Concepts: Preformation vs Epigenesis
• The morphogenetic process called gastrulation
• The question of how a zygote becomes an animal has rearranges the cells of a blastula into a three layered
been asked for centuries. (triploblastic) embryo, called a gastrula, that has a
• In the 18th century, the prevailing theory was a notion primitive gut.
called preformation – the idea that the egg or sperm o Diploblastic organisms have two germ
contains an embryo. layers
o A preformed miniature infant, or • The three tissue layers produced by gastrulation are
“homunculus,” that simply becomes larger called embryonic germ layers.
during development. o The ectoderm forms the outer layer of the
• Kaspar Friederich Wolff (1759) demonstrated there gastrula.
was no preformed chick in the early egg. ▪ Outer surfaces, neural tissue
o Undifferentiated granular material became o The endoderm lines the embryonic digestive
arranged into layers. tract.
o The layers thickened, thinned, and folded to o The mesoderm partly fills the space
produce the embryo. between the endoderm and ectoderm.
• Epigenesis is the concept that the fertilized egg
contains building materials only, somehow assembled Building a Body Plan
by an unknown directing force.
• Cell differentiation – the specialization of cells in
o Although current ideas of development are
their structure and function.
essentially epigenetic in concept, far more is
• Morphogenesis – the process by which an animal
known about what directs growth and
takes shape and differentiated cells end up in their
differentiation
appropriate locations.
• The sequence includes: Animal Body Plans
o Cell movement
o Changes in adhesion • Body plans are constrained by ancestry, major
o Cell proliferation features may become modified, but are rarely lost.
• There is no “hard-wired” master control panel • Animal body plans differ in their grade of organization,
directing development. body symmetry, number of germ layers, and type of
o Sequence of local patterns in which one step body cavity.
in development is a subunit of another.
Symmetry
o Each step in the developmental hierarchy is
a necessary preliminary for the next. • Spherical Symmetry - occurs when any plane
passing through the center divides the body into
Organogenesis
mirror image halves.
• Various regions of the three embryonic germ layers o Mostly found among the protists.
develop into the rudiments of organs during the • Radial Symmetry - applies when more than two
process of organogenesis. planes passing through the longitudinal axis can
• Many different structures are derived from the three divide the organism into mirror image halves.
embryonic germ layers during organogenesis o Jellyfish
• Biradial Symmetry - two planes will divide the
TOPIC 07: ARCHITECTURAL PATTERN OF AN ANIMAL organism.
o Comb jellies
The Appearance of Major Body Plans • Bilateral Symmetry is found in organisms where one
plane can pass through the organism dividing it into
• The Cambrian explosion marks the earliest fossil right and left halves.
appearance of all major groups of living animals plus o Better for directional movement.
some groups that are only known from fossils. o Monophyletic group called Bilateria.
o All major body plans appeared at this time.
▪ Result of extensive selection. Other Key Features of Body Plans
▪ Are limiting determinant of future
adaptational variants. • Segmentation is a serial repetition of similar body
segments along the body.
Body Plans o Each segment is a metamere or somite.
o May include external & internal components.
• One way in which zoologists categorize the diversity o Obscured in many animals, like humans.
of animals is according to general features of o Permits greater body mobility and complexity
morphology and development. of structure & function.
• A group of animal species that share the same level
of organizational complexity is known as a grade. Components of Metazoan Bodies
Connective Tissue
Muscle Tissue
Nervous Tissue
Speciation
• Speciation = formation of a new species
Binomial Nomenclature • Happens when a population becomes different
enough that they can no longer reproduce
• Is the system Linnaeus used for naming species. successfully with the original group
o Genus and species
• Names are latinized and italicized, only the genus is The Typological Species Concept
capitalized.
• The morphological species concept defines species
o Ex. Common Name – Banana;
based on their physical form and structure.
Scientific Name – Musa Paradisicum
• Cons - can be misleading, subjective
o Ex. Common Name – Cat
Scientific Name – Felis Catus Typological (Morphological) Species Concept
Species • Classifies organisms based on physical appearance
and structure.
• Defining a species can be difficult.
• If organisms look similar, they are considered the
• Can you recall the different methods to infer
same species.
evolutionary relationship?
• Criteria: Cons: Can be misleading because some organisms may look
o Common descent alike but are actually different species.
o The smallest distinct groupings of organisms
sharing a pattern of descent. The Biological Species Concept
▪ Morphological & molecular
techniques Members of a species • A species is a group of members who can interbreed
must form a reproductive and produce viable, fertile offspring.
community that excludes other • “A species is a reproductive community of populations
species. (reproductively isolated from others) that occupies a
specific niche in nature.” Mayr 1982
Biomolecules / Macromolecules in Evolution • Cons - inapplicable to asexual and extinct organisms
• Biomolecules (macromolecules) like DNA,
Biological Species Concept
proteins, and RNA are used to:
o Compare organisms at the molecular level • Classifies organisms based on their ability to
o Infer evolutionary relationships interbreed and produce fertile offspring.
• This is more accurate than just physical • If they can successfully reproduce, they belong to the
appearance because it shows genetic similarity. same species.
• A phylogenetic tree is one way of representing • A type of phylogenetic tree that focuses on the order
evolutionary relationships. of branching based on shared derived characteristics,
o A phylogenetic tree is a branching diagram without necessarily indicating the amount of
that illustrates evolutionary relationships. evolutionary change.
o Branches represent evolutionary lineages.
o Nodes represent common ancestors Major Divisions of Life
• Phylogenies are inferred by identifying organismal
Kingdom and Domains
features, characters, that vary among species.
Bacteria Archaea Eukarya
o Morphological → physical structures
The Six-Kingdom System
o Chromosomal → chromosome similarities
o Molecular → DNA, RNA, proteins Bacteria Archaea Protista Plantae Fungi Animalia
o Behavioral or ecological → behavior and The Traditional Five-Kingdom System
environment Monera Protista Plantae Fungi Animalia
Example: Mitochondria
• A big cell consumed a small bacteria that could
make energy.
• It stayed alive inside the cell.
• Later, it became the mitochondria.
o Gives energy to the cell
o Has its own membrane and DNA
Chloroplast
• Some cells later consumed photosynthetic
bacteria.
• They were not digested.
• These became chloroplasts.
o helps plants make food using sunlight
(photosynthesis)
TOPIC 02: UNICELLULAR EUKARYOTES
Think of it like this:
Emergence of Eukaryotes • Big cell = house
• Small cell = helper
• First evidence of life dates to 3.5 billion years ago.
The helper stayed inside the house and worked there
o First cells were bacteria like.
permanently.
o Endosymbiotic Theory
• Origin of complex eukaryote cells Term Meaning
o Most likely symbiosis among prokaryotic Prokaryote Simple cell without nucleus
cells. Eukaryote Complex cell with nucleus
Symbiosis Two organisms living together and helping
First Evidence of Life each other
• The first evidence of life appeared around 3.5 Endosymbiosis One organism living inside another organism
billion years ago. Mitochondria Organelle that produces energy
Chloroplast Organelle used for photosynthesis
• The earliest living things were simple bacteria-like
cells, also called prokaryotic cells. Easy Summary
• Prokaryotes are simple cells that do not have a
nucleus and do not have membrane-bound • Life started with simple bacteria-like cells.
organelles. • Prokaryotic cells lived first before eukaryotic cells.
• Some large cells consumed smaller cells but did not
What are Eukaryotes? digest them.
• Eukaryotic cells are more complex cells. • The smaller cells lived inside and helped the larger
• They have: cells survive.
o a nucleus • Over time, they became organelles like:
o membrane-bound organelles like o mitochondria
mitochondria and chloroplasts o chloroplasts
• Examples of eukaryotes: • This idea is called the Endosymbiotic Theory.
o plants
o animals Kingdom Protista
o fungi
o protists • Protists are simple eukaryotic organisms that do
not belong to the categories of plants, animals, or
Endosymbiotic Theory; explains how complex eukaryotic fungi. While they are primarily unicellular, some can
cells formed from simple prokaryotic cells. exist as colonies of cells.
• They are commonly found in aquatic environments,
Simple Explanation: moist terrestrial habitats, or as parasites
1. A bigger prokaryotic cell ate a smaller prokaryotic • All (protists) are eukaryotic.
cell. o This means their cells have:
2. The smaller cell was not digested. ▪ a nucleus
3. The two cells lived together and helped each other ▪ membrane-bound organelles
survive – Symbiotic Relationship o Protists = eukaryotic
4. Over time, the smaller cell became part of the o Bacteria = prokaryotic
bigger cell. • All have cells with nuclei.
o A nucleus is the control center of the cell.
This process helped form important organelles inside o It stores the cell’s DNA or genetic material.
eukaryotic cells. o This is one reason protists are more complex
than bacteria.
• Can be unicellular or multicellular.
o Many protists are unicellular (only one cell). • Fungus Like
o Some are multicellular and made of many o They are similar to fungi because they:
cells. ▪ absorb nutrients from dead
▪ Example: materials
▪ Amoeba = unicellular ▪ usually live in moist places
▪ Giant algae/seaweed = o Most help break down dead organisms.
multicellular
• Can be microscopic or over 100 meters (300 feet) Basis of Classification of Protists
long.
Protists are classified based on:
o Some protists are so small they can only be
seen with a microscope. • mode of nutrition (how they get food)
o Others, like some seaweeds, can grow very
• presence of cell wall (if they have a cell wall or not)
large.
• Some are heterotrophs, while others are Type of Protist Similar To How They Get Food
autotrophs. Protozoa Animals Eat other organisms
True Algae Plants Make food through photosynthesis
o Heterotrophs; Amoeba get food by eating
Fungus-like Protists Fungi Absorb nutrients from dead matter
or absorbing nutrients from other organisms.
o Autotrophs; Algae make their own food
Protozoans (Animal like Protista)
using sunlight through photosynthesis.
• Protozoans are an extremely diverse assortment of
Protists are simple eukaryotic organisms that do not
unicellular eukaryotes
belong to plants, animals, or fungi.
o Protozoans are a very varied group of
single-celled organisms with a nucleus.
• Protists are organisms with a nucleus and
complex cells. Protozoans
• They are grouped separately because they are
different from plants, animals, and fungi. • Lacks a cell wall
• Have at least one motile stage in the life cycle
• Most protists are single-celled (unicellular).
• Most ingest their food
• Some live in colonies, where many cells stay
• Carry on all life activities within a single cell.
together but still function independently.
• Can survive only within narrow environmental ranges.
Example: Amoeba, Paramecium and Algae • Very important ecologically
Entamoebas
3. Parasitic – one benefits at a cost to the other
o One organism benefits, while the other is • Entamoebas are parasites of vertebrates and some
harmed. invertebrates.
o Ex. Plasmodium (protozoan) causes malaria • Entamoeba histolytica causes amoebic dysentery
in humans. in humans
Categories Based on Locomotion o Are parasites that live inside animals
(vertebrates and some invertebrates)
• Flagellates o Can cause diseases in humans
o Move using a long whip-like tail called a o Entamoeba histolytica → causes amoebic
flagellum. dysentery (diarrhea, stomach pain)
o Ex:
Form & Function
▪ Euglena
▪ Trypanosoma (causes sleeping 1. Lobopodia
sickness) o Shape: Broad, blunt, finger like.
▪ Thick, finger-like projections
• Pseudopod (Sarcodines / Amoeboids) ▪ Used for movement and eating
o Move using false feet called pseudopodia 2. Filopodia
(temporary body extensions). o Shape: Slender, thread like, and pointed.
o Ex: Amoeba ▪ Thin, thread-like, pointed
extensions
• Ciliates ▪ Used for sensing and movement
o Move using tiny hair-like structures called 3. Reticulopodia
cilia. o Net like or branched, forming a complex
o Ex: Paramecium network.
▪ Branching, net-like structures
• So, the particular basis is their locomotory ▪ Used to trap food
structure. 4. Axopodia
o Shape Long, thin, stiff, and radiating like
Locomotion in Protozoan
spokes of a wheel
Protists have three main methods of locomotion. ▪ Thin, sharp, ray-like (spoke of a
wheel)
• Flagellates - Some protists, like this Giardia and ▪ Used for support and capturing food
Trypanosoma.
o Move using a long whip-like tail
(flagellum).
• Fission is the cell multiplication process in protozoa. 1. Syngamy – gametes from two individuals fuse to
o A method where a protozoan reproduces form the zygote.
by dividing its cell to form new organisms. o Two protozoans fully join and form a new
organism.
1. Binary Fission – one individual splits into two equal o Closest form to sexual reproduction.
sized individuals.
o One cell divides into two equal-sized 2. Autogamy – gametes from one individual fuse.
cells. o A single protozoan uses its own genetic
material to reproduce.
2. Budding – progeny cell is much smaller than parent. o No partner needed.
o A new organism forms as a small
outgrowth of the parent cell. 3. Conjugation – No new cells are formed, but genetic
o The new one is smaller than the parent. recombination occurs.
o Two protozoans come close and exchange
3. Multiple Fission – multiple nuclear divisions followed genetic material.
by multiple cytoplasmic divisions producing several o They do not fully merge, just share genes
offspring for variation.
o The nucleus divides many times first, then
the cell splits into many new individuals. Simple Summary
o Produces several offspring at once.
• Binary fission → 1 becomes 2
• All of above accompanied by some form of mitosis. • Multiple fission → 1 becomes many
o Mitosis in protozoa divisions varies from • Syngamy → full fusion
metazoan mitosis. • Autogamy → self-reproduction
o All protozoan cell divisions happen with • Conjugation → exchange genes without merging
some form of mitosis (cell division
Cysts
process).
o But protozoan mitosis is different from • Many protists are able to survive harsh conditions
animals (metazoans). through the formation of cysts, dormant forms with
▪ Nuclear membrane often persists. resistant outer coverings and a shutdown of
▪ Spindle may form within the nuclear metabolism.
membrane. o Some protists can survive harsh or
▪ Centrioles not observed in ciliates. unfavorable conditions by forming cysts.
▪ Macronucleus of ciliates elongates, o A cyst is a resting (dormant) stage.
constricts, and divides without
mitosis (amitosis) What happens in a cyst?
Differences in Protozoan Mitosis • The cell forms a thick protective outer covering.
• The cell’s activity slows down or stops (low
• The nuclear membrane may stay intact during
metabolism).
division.
• It can survive until conditions become good again.
• The spindle fibers can form inside the nucleus.
• In ciliates, centrioles are not seen. Simple idea: Cyst = “sleep mode” of protists to survive
extreme conditions.
Special Case in Ciliates
Plant-Like Protist (Algae)
• The macronucleus divides in a simpler way.
• It just stretches, narrows, and splits. • Plant-like protists, also known as algae, are
• This is called amitosis (no typical mitosis process). organisms that photosynthesize like plants but lack
some of their more complex features.
Many types of protists reproduce sexually as well as • Each has chlorophyll and other photosynthetic
asexually. pigments
o Plant-like protists are called algae.
Types: o They can make their own food using
sunlight (photosynthesis).
1. Isogametes – Fusion of morphologically similar o They contain chlorophyll and other
gametes pigments, which give them different colors.
o Two gametes that are same in shape and o They are simpler than true plants.
size join together.
Origin of Metazoa
• The skeletal framework of a sponge may be fibrous • Sponges have remarkable regeneration capabilities.
or rigid. o Sponges can regrow lost parts easily
o Sponges have an internal support structure (regeneration).
(skeleton). • Regeneration following fragmentation is a form of
• Fibrous Skeleton as composed mainly of Spongin asexual reproduction.
(elastic and soft) o If a sponge breaks into pieces, each piece
o Soft - mainly compose of sponging can grow into a new sponge (asexual
o Sponging - is elastic and soft reproduction).
• Rigid Skeletons consist of needlelike spicules. • External Buds can break off to form new sponges.
o Calcareous → made of calcium o Small outgrowths (buds) form on the
o Siliceous → made of silica (glass-like) sponge surface.
• Composition and shape the spicules forms the basis o These buds can detach and become new
of sponge classification. sponges.
o The type and shape of spicules are used • Internal Buds (gemmules) in freshwater sponges
to classify different sponges. can remain dormant in times of drought
o Found in freshwater sponges.
Canal Systems
o These are internal survival structures.
• Asconoid – the simplest canal system. o They can stay inactive (dormant) during
o Choanocytes line the spongocoel. harsh conditions like drought and later grow
o Water enters through the ostia and exit into a new sponge.
through the large osculum.
o Usually tube shaped. • Most sponges are Hermaphrodites meaning that
o Found only in the Class Calcarea. each individual functions as both male and female.
o meaning one sponge can produce both
Asconoid (Simplest) sperm and eggs.
• Water goes straight into a central cavity. o The sex cells (gametes) are derived from
• Choanocytes line the main cavity (spongocoel). choanocytes or sometimes archeocytes.
• Water enters through ostia and exits through
• Most sponges are viviparous.
osculum.
o Most sponges are viviparous, meaning the
• Body is usually tube-shaped. young are born as larvae instead of being
• Found in Class Calcarea. released as eggs
For example:
REST
• Corals and hydras are usually sessile and exist in
R – radial the polyp form.
E – endoderm and ectoderm • Jellyfish are free-swimming and exist mostly in the
S – sting medusa form.
T – tentacles
• Polymorphism some species exist as both polyps
Radiate Animals and medusae during their life cycles.
o which means one species can have more
• Phylum Cnidaria than one body form during its life cycle.
o Radial Symmetry - their body parts are
arranged around a central point. A cnidarian may start as a polyp (attached form), then later
o Diploblastic – 2 embryonic tissue layers develop into a medusa (swimming form). Each form has a
▪ meaning they develop from two different function:
embryonic tissue layers.
• Polyp → feeding and asexual reproduction
• All animals except sponges belong to the clade • Medusa → movement and sexual reproduction
Eumetazoa, the animals with true tissues. This alternation of forms helps cnidarians survive and
o All animals except sponges belong to this reproduce more effectively in aquatic environments.
group because sponges lack organized
tissues.
Cnidarians have:
• Radial symmetry
• Tissues – derived from two embryonic germ layers
Polyp • Cnidocytes – special cells with stinging organelles
called nematocysts
• Tube-shaped body form o Their tentacles contain cnidocytes, special
• Usually attached to a surface stinging cells with nematocysts used for
• Mouth faces upward capturing prey and defense.
• Example: Sea anemones, corals • Gastrovascular cavity – extracellular digestion
• Tentacles have cells called cnidocytes
Medusa • Digestion of Cnidaria: Incomplete
o They also have a gastrovascular cavity for
• Bell or umbrella-shaped body form
digestion. Digestion is incomplete because
• Free-swimming
they only have one opening used for both
• Mouth faces downward food intake and waste removal.
• Example: Jellyfish
o Finished inside gastrodermis cells
o Digestion is both extracellular and
intracellular
Reproduction
Body Wall
Class Cubozoa
Class Anthozoa
• Phylum Mollusca includes snails and slugs, oysters When present, the calcareous shell is secreted by the mantle
and clams, octopuses and squids. and is lined by it. It has 3 layers:
Common Characteristic: They all have a soft body • Periostracum - the outer organic layer helps to
("mollusks" means soft). protect the inner layers from boring organisms.
• Prismatic Layer - Middle, strong, and structural
Mollis – simply the Latin word for "soft." layer.
• Nacreous Layer - Inner, and smooth - surrounds
• Gastropod means "stomach footed" referencing the foreign objects to form pearls in some.
position of the animals "foot" beneath its gut
o Their "foot" is located right under their Mantle Cavity
belly/stomach area.
o Examples: Snails and slugs. • The mantle cavity is the space between the mantle
• A bivalve's name references its two shells or and the visceral mass.
halves, called valves o The respiratory organs (gills or lungs) are
o They have a shell made of two hinged parts generally housed here.
that open and close.
Mollusk Life Cycle / Reproduction
o Examples: Clams, oysters, and mussels.
• Cephalopod means "head footed" referencing the • Most mollusks reproduce sexually, with separate
way a cephalopods' head connects to its many arms sexes (dioecious), though some are
o Their head is directly connected to their hermaphroditic.
"foot" (which is divided into tentacles). • Fertilization may be external (e.g., bivalves) or
o Examples: Octopus, squid, and cuttlefish. internal (e.g., cephalopods and some gastropods).
• The life cycle of many mollusks includes a free-
Phylum Mollusca
swimming, ciliated larval stage called a
• Mollusks evolved in the sea, and most mollusks are trochophore.
still marine. • The trochophore larval stage is followed by a free-
o Some gastropods and bivalves inhabit swimming veliger larva in most species.
freshwater.
Mollusks Life Cycle:
o A few gastropods (slugs & snails) are
terrestrial. 1. Trochophore Larva: First 6–12 hours.
o The egg hatches into a tiny, translucent larva
Humans & Mollusks
with a ring of fine hairs (cilia) used for
• A few are pests or introduced nuisances: swimming.
o Shipworms - burrow through wood, including 2. Veliger Larva: Next Stage.
docks & ships. o The trochophore develops further, starting to
o Terrestrial snails and slugs damage garden grow a tiny shell and a swimming device
plants. called a velum.
o Mollusks serve as intermediate hosts for 3. Pediveliger Larva: Settling Stage.
many parasites. o The larva grows a muscular foot (indicated
by the prefix pedi-). It uses this foot to find a
Mollusks Body Plan (Squidward) – moody, very, miserable permanent spot to settle down.
4. Metamorphosis: Final Transformation.
• Muscular Foot o The larva undergoes a major change to look
• Visceral Mass - containing circulatory, respiratory exactly like a mini version of the adult,
and reproductive organs. dropping its swimming parts.
• Mantle houses the gills and, in some cases, secretes
a protective shell over the visceral mass. Note on Variations:
• Most gastropods are herbivores and feed by TOPIC 06: PHYLUM ECHINODERMATA
scraping algae off hard surfaces using the radula.
• The Greek words for the phylum Echinodermata are
• Some are scavengers of dead organisms, again ”Echinos” which means "spiny", and Derma, which
tearing off pieces with radular teeth. means "skin".
• Some are carnivores and have a radula modified into • Echinoderms include sea stars, brittle stars, sea
a drill to bore through the shells of other mollusks. urchins, and sea cucumbers.
They use chemicals to soften the shell. • Entirely marine (they can only survive in salt water or
Class Bivalvia marine water)
• Lack of ability to osmo-regulate (ex. pag nag
• Bivalved mollusks have two shells (valves). increase yung salt content of humans there is a
• Mussels, clams, oysters, scallops, shipworms. tendency na baba yan)
• Non-segmented.
• Mostly sessile filter feeders.
• No head or radula. Body Forms (Patrick)
• Part of the mantle is modified to form incurrent and
excurrent siphons. Mnemonic: Patrick, Star, Wears, Designer, Pants (PSWDP)
o Used to pump water through the organism
• Pedicellariae
for gas exchange and filter feeding.
• Spiny endoskeleton
o Sometimes used for jet propulsion.
• Water vascular system
Class Bivalvia – Locomotion • Dermal branchiae (skin gills)
• Pentaradial symmetryin adults
• Bivalves move around by extending the muscular foot o 5 arms are arranged in circle manner
between the shells.
• Scallops and file shells swim by clapping their shells Symmetry
together to create jet propulsion.
• Echinoderms are bilaterally symmetrical as larvae.
Class Cephalopoda – Locomotion o This means their ancestors were bilaterally
symmetrical.
• Cephalopods include octopuses, squid, nautiluses, • As adults they show secondary radial symmetry –
and cuttlefish. pentaradial (5 parts)
• Marine carnivores with beak-like jaws surrounded by • Today’s echinoderms are mostly motile.
tentacles of their modified foot. • Many are still radial.
• The modified foot is a funnel for expelling water from • Some have again become superficially bilateral
the mantle cavity. (skeletal & organ systems still pentaradial).
• Sea cucumbers.
• No well-defined head or brain.
Water Vascular System • They can also cast off an injured arm.
• Some species can even regenerate an entire
• Echinoderms have water vascular system. individual from a broken off arm.
• A system of canals and specialized tube feet that • Usually, a small piece of the central disc must be
functions in: included.
o Locomotion • Linckia can regenerate a whole new individual from a
o Food gathering broken arm with no central disc attached.
o Respiration
o Excretion Class Echinoidea
• The water vascular system opens to the outside
through small pores in the madreporite. • Class Echinoidea includes sea urchins and sand
o Serves as entry point of the water, yung dollars
tubig papasok siya sa madreporite. • The endoskeleton is well developed in echinoids.
• Madreporite (Sieve plate) • Dermal ossicles have become close-fitting plates.
• Radial canal • Echinoids lack arms, but still show the pentamerous
• Ampulla plan in the five ambulacral areas with pores in the test
• Tube Foot (Movement) for the tube feet.
• Some urchins have very reduced tests, and bright
Locomotion: Water Vascular System coloration.
• The pedicellariae in these species contain painful
Endoskeleton
toxins.
• Echinoderms have an endoskeleton of calcareous • Sand dollars use their short spines to move sand
ossicles, often with spines. & its organic contents to the sides, the food particles
drop between the spines, and ciliated tracts on the
Major Classes oral side carry the particles to the mouth.
Class Holothuroidea
Subphylum Trilobita (Tri means 3) • Three body regions: head, thorax, abdomen
• 1 pair of antennae
They have 3 lobes ang kanilang cephalon, thorax, pygidium. • 3 pairs of legs (6 total)
• Most have wings (usually 2 pairs)
• Extinct marine arthropods
• Compound eyes
• Extinct animals na sila - Dominated oceans 521
• Dominated by the Class Insecta
million years ago (Paleozoic Era)
o They have no longer descendants • Found in nearly every habitat on Earth
• Flattened, segmented body with 3 lobes (tri-lobed) Comparing the Subphyla
• 1 pair of antennae, compound eyes
• Likely bottom-dwellers or scavengers
• No modern descendants
Subphylum Chelicerata
Chelicerata (Fang-Bearers)
Subphylum Myriapoda (myria means many, poda means Evolution And Adaptations
legs)
• Evolved from segmented ancestors (trilobites) with
Many-Legged Wanderers hardened exoskeletons
• Extensive fossil record, including trilobites
• Elongated bodies with many similar segments • Adaptations like flight, complex behaviors, and
o Mahaba yung body nila diverse feeding habits
• 1 pair of antennae
• Simple eyes (or none) Nutrition in Arthropods
• Terrestrial only
• Varied diets: herbivores, carnivores, omnivores,
• Two main groups: parasites
• Centipedes (Chilopoda) – 1 pair of legs/segment, • Specialized mouthparts for feeding
fast predators, venomous
• Filter feeding (crustaceans), predation (spiders),
o Presence ng legs per segment
scavenging (insects)
Locomotion in Arthropods • Compound eyes and advanced chemoreceptors aid in
environmental perception
• Jointed appendages adapted for walking, swimming,
jumping, or flying Reproductive Strategies in Arthropods
• Muscles attached to exoskeleton for movement
• Insects: wings for flight, highly efficient locomotion • Mostly dioecious with internal fertilization, but some
• Crustaceans: swimmerets and walking legs exhibit external fertilization (e.g., crustaceans)
o Mag kaiba yung male and female repro
Digestion in Arthropods • Metamorphosis is common in insects:
holometabolous (complete) vs. hemimetabolous
• Complete digestive system with specialized regions (incomplete)
• Foregut (ingestion & storage), midgut (digestion & • Some arthropods (e.g., aphids) exhibit
absorption), hindgut (waste processing) parthenogenesis
• Enzymatic digestion varies among different groups • Complex mating behaviors in spiders and
crustaceans (e.g., courtship rituals, pheromone
Reproduction in Arthropods
signaling
• Mostly sexual reproduction, some parthenogenesis
TOPIC 08: THE WORMS
• Internal or external fertilization
• Metamorphosis common in insects (complete and
Worms
incomplete)
o Complete - development with four stages: • Phylum Platyhelminthes – flat body
egg, larva, pupa, and adult. • Phylum Nematoda – round worms
o Incomplete metamorphosis - development • Phylum Annelida – body have segments
where the young looks like a small adult and
gradually grows without a pupal stage. Introduction
• Complex mating behaviors and parental care in some
species • Bilateral symmetry - the simplest organisms to
exhibit this type of symmetry
• Invertebrates - lack backbone
• Made of the three germ layers: endoderm,
ectoderm, and mesoderm.
o Triploblastic
• Undergo cephalization
• Platyhelminthes - General Features
• Commonly known as flatworms
• Unique Features:
Main difference ng nymphal stages sa metamorphosis o 1. Dorso-Ventrally Flattened Body,
• Nymphal stage: hatching stage, parang lumalaki o 2. Acoelomate,
lang siya o 3. Triploblastic
• Metamorphosis: From the fertilized egg is very • Examples: Planaria, Tapeworms, Liver Flukes
different from the hatched one Platyhelminthes - Nutrition
Advanced Nutrition in Arthropods • Free-living species (e.g., Planaria): Feed on small
• Highly diversified feeding strategies: predation, organisms through a pharynx
herbivory, detritivory, parasitism • Parasitic species (e.g., Tapeworms): Absorb
• Adaptations include specialized mouthparts nutrients from the host's intestine
(mandibles, chelicerae, proboscis) • Lack a complete digestive system, only a
• Crustaceans use filter feeding mechanisms; gastrovascular cavity (Incomplete)
arachnids use external digestion via enzymatic Platyhelminthes - Locomotion
breakdown
• Some insects, like termites, have symbiotic gut • Cilia in free-living species for gliding (planaria)
bacteria for cellulose digestion • Muscular contractions allow movement in absence
of cilia
Morphological Specialization in Arthropods
• Parasitic species have reduced locomotion and
• Body segmentation (metamerism) with rely on host movement
tagmatization (fusion of segments for specialized Platyhelminthes - Reproduction
functions)
• Exoskeleton composed of protein and chitin provides • Asexual reproduction by fission and regeneration
defense and prevents desiccation (Planaria can regrow entire bodies)
• Molting (ecdysis process) regulated by ecdysteroid • Sexual reproduction: Hermaphroditic but usually
hormones allows for growth. undergo cross-fertilization.
o Pinaka mode pa rin ng reproduction nila is Comparative Table
sexual reproduction
• Platyhelminthes:
Nematoda - General Features o Acoelomate
o Incomplete Digestion
• Commonly known as roundworms o Cilia/Muscles
• Unique Features: o Asexual & Sexual
o 1. Pseudocoelomate, • Nematoda:
o 2. Cylindrical body covered with tough cuticle o Pseudocoelomate
o Complete Digestion
• Examples: Ascaris (female) and Hookworms. o Longitudinal Muscles
• Found in diverse environments, from soil to o Sexual
parasites in animals • Annelida:
o Coelomate
Nematoda - Nutrition
o Complete Digestion
• Free-living species: Decomposers feeding on o Setae
bacteria, fungi, and detritus o Mostly Sexual
• Parasitic species:
TOPIC 09: CHORDATES
o Hookworms suck blood,
o Ascaris absorbs digested food from hosts
Phylum Chordata
• Complete digestive system with a separate mouth
and anus • By the end of the Cambrian period, 540 million years
ago, an astonishing variety of animals inhabited
Nematoda - Locomotion
Earth’s oceans.
• Hydrostatic skeleton (fluid-filled pseudocoelom) • One of these types of animals gave rise to
aids movement vertebrates, one of the most successful groups of
• Thrashing, whip-like motion animals
• Chordates are bilaterian animals that belong to the
Nematoda - Reproduction clade of animals known as Deuterostomia.
• Five distinctive characteristics define the
• Sexual reproduction: Most are dioecious (separate chordates:
male and female) o Notochord
• Internal fertilization with copulation o Dorsal tubular nerve cord
• Females can lay thousands of eggs per day o Pharyngeal pouches (gill slits)
o Endostyle
Annelida - General Features
o Postanal tail
• Commonly known as segmented worms • All are found at least at some embryonic stage in all
• Unique Features: chordates, although they may later be lost.
o 1. Metamerism (Segmentation),
Notochord
o 2. True Coelom,
o 3. Closed Circulatory System • The notochord is a flexible, rod-like structure derived
• Examples: Earthworms, Leeches, from Mesoderm.
• Segmentation allows specialization of body regions o The first part of the endoskeleton to appear
in an embryo.
Annelida - Nutrition o Place for muscle attachment.
o In vertebrates, the notochord is replaced by
• Earthworms: Eat decaying organic matter, crucial
the vertebrae.
for soil aeration
▪ Remains of the notochord may
• Leeches: Ectoparasites feeding on blood
persist between the vertebrae.
(anticoagulant enzymes)
Dorsal Tubular Nerve Cord
Annelida - Locomotion
• In chordates, the nerve cord is dorsal to the
• Earthworms: Move using setae and peristaltic
alimentary canal and is a tube.
motion
o The anterior end becomes enlarged to form
• Leeches: Use suckers for inchworm-like
the brain.
movement
o The hollow cord is produced by the infolding
Annelida - Reproduction of ectodermal cells that are in contact with
the mesoderm in the embryo.
• Earthworms & Leeches: Hermaphroditic (may egg o Protected by the vertebral column in
and sperm in one), cross-fertilization via clitellum vertebrates.
• Some reproduce by budding or fragmentation
Pharyngeal Pouches and Slits • Includes fishes, amphibians, reptiles, birds, and
mammals.
Pharyngeal slits are openings that lead from the pharyngeal • Divided into Agnatha (jawless), Pisces (fishes), and
cavity to the outside. They are formed when pharyngeal Tetrapoda (four limbed vertebrates).
grooves and pharyngeal pouches meet to form an opening.
Key Features - Vertebrata
• The perforated pharynx evolved as a filter feeding
apparatus. • Closed circulatory system with a heart (2-4
• Later, they were modified into internal gills used for chambers).
respiration. • Paired appendages
o (fins in fishes, limbs in tetrapods).
Endostyle or Thyroid Gland
• Respiration: Gills in aquatic species, lungs in
• The endostyle in the pharyngeal floor, secretes terrestrial species.
mucus that traps food particles. Key Features - Agnatha (Jawless Vertebrates)
o Found in protochordates and lamprey larvae.
o Secretes iodinated proteins. • Lack jaws and paired fins.
• Homologous to the iodinated-hormone-secreting • Cartilaginous skeleton, no vertebral column
thyroid gland in adult lampreys and other vertebrates • Notochord Retained
• Slimy, scaleless body.
Postanal Tail
• Eel-like Body Shape.
• The postanal tail, along with somatic musculature and
Agnatha – Nutrition
the stiffening notochord, provides motility in larval
tunicates and amphioxus. • Ectoparasitic or scavengers.
o Evolved for propulsion in water. • Nutrition: Feed on blood or decaying matter.
o Reduced to the coccyx (tail bone) in
• Lampreys (parasitic or filter-feeding) and hagfish
humans.
(scavengers that produce slime for defense).
Classification of Chordata • Simple Digestive System
• Nutrients are absorbed directly as food passes
through their digestive tract.
Agnatha – Reproduction
Filter Feeders – Use gill rakers to trap plankton (e.g., whale • Larval Stage (Tadpoles) – Mostly Herbivorous or
sharks, manta rays). Omnivorous-Feed on algae, plant material, and
microorganisms.
• Suction Feeders – Expand their mouth to create a • Adult Stage – Mostly Carnivorous
vacuum (e.g., groupers). o Feed on insects, worms, spiders, small fish,
• Biting/Grasping Feeders – Have strong jaws and and other invertebrates.
teeth to bite prey (e.g., barracuda). • Some large amphibians eat small vertebrates (e.g.,
Pisces - Reproduction snakes, birds, rodents).